This means the shape is determined by its function, or the the combination of parts that create the function. Its a saying or way of working originally applied to designing products or buildings , but it works for anything you want to design and is a good way of choosing or deciding the success of designs. It sounds a bit dry and featureless- sort of no frills, but it does incorporate decoration and styling for example if that serves the function of what it is you are designing.
Natural things and living things are almost the ultimate example of things refined for their purposes of the organism, although not exactly designed, it aplies even the most extravagant and flamboyant looking creatures or plants.
Natural things often behave or function in a way to minimisation of energy expenditure, or make it as efficient as possible.
Seacreature-shark fast moving/efficient- streamlined to reduce drag- VS weedy sea dragon- doesn't have to move fast- camouflage of ornate fins looks fanciful but help it hide.
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